Professor Emeritus
EBIO
Plant physiological ecology; forest-atmosphere exchanges of carbon dioxide, water vapor, ozone and energy; environmental and physiological control over the emission of isoprene and other biogenic volatile organic compounds (VOCs) from forest ecosystems and the relationship of VOC emissions to atmospheric chemistry; coupling of biogeochemical cycles, especially those involving water, carbon and nitrogen; the evolution of C4 photosynthesis in plants.