Topher received a BA from Earlham College in May 2010 and majored in Biology. Topher completed his PhD in the Department of Ecology and Evolutionary Biology and was a member of Brett Melbourne’s lab. He did his lab rotations in Dr. Rob Knight’s lab in Biochemistry, in Dr. Manuel Lladser's lab in Applied Mathematics and in Dr. Brett Melbourne's lab in Ecology and Evolutionary Biology. Topher graduated in Spring 2017.