Topic 50. The response of the eye to color

 

          The motivation for the previous discussions in which we tried to match any color using various combinations of three primary colors is that this process is based on how the eye really perceives color.

 

          The rods of the eye – the sensors that are used when the light intensity is low, have some variation in their response to different colors. They are most sensitive to colors in the green portion of the spectrum near 500 nm, so that when the light is dim it is easiest to see objects that either radiate or reflect green light. However, since there is only one type of rod sensor, the rods cannot be used to discriminate among different colors, since they respond only to the weighted sum of the intensity that strikes them (where wavelengths in the vicinity of 500 nm green are more heavily weighted than those in the red or blue). 

 

The cones of the eye – the sensors that are used when the light intensity is higher, come in three different types, where the differences among the types are the wavelength bands they respond to most strongly.  The three types of cones respond to all wavelengths to some extent, so that every color excites them all – at least to some extent. However, the amplitude of the response of each type is a function of the incident wavelength.

 

Unfortunately, each type of cone does not provide any wavelength discrimination beyond its overall response function. In other words, the short-wavelength cone, which is sensitive primarily to a broad band of colors in the blue and green portions of the spectrum, cannot provide any information on the wavelength of the incident light beyond the fact that it has been triggered by some wavelength within its response band. Color discrimination depends on the fact that every wavelength excites the three types of cones by different amounts – every incident wavelength produces a unique triplet of excitation values. This is an important design feature: if a band of wavelengths excited only one type of the cones, then we would not be able to distinguish differences in hue within this band, and if the response functions did not overlap, then there would be wavelengths that we would not be able to see at all, since those wavelengths would not excite any one of the cones or would produce so little excitation that these wavelengths would appear very dim. Finally, if the response functions of the cones were essentially identical over any range of wavelengths then it would be difficult or impossible to distinguish the hues in this wavelength range.

 

The details of the response functions of the 3 types of cones can be derived from the observed response of the eye to various colors – especially to the fact that only two complementary colors can combine to produce white. These pairs of colors must therefore straddle the cross-over points between the sensitivities of the 3 cones, so that the two colors can excite all 3 types of cones more or less equally so as to give the sensation of white.

 

Additional information comes from people who are “color blind.” These people often have only two of the 3 types of cones, so that every color that they see is some combination of the two types of cones that are working. For example, if the green cones are not working, then these people discriminate among colors based on the red and blue content alone, and they cannot distinguish colors that have the same red and blue content but which differ only in how much green they contain. In other words, they see all colors as if they were on vertical lines through the color horseshoe (figure 9.11, page 245), since colors along these lines differ only in how much green they contain.

 

The result of these considerations is that the response curves of the 3 types of cones are as shown in the following figure, which is taken from the textbook on page 273 (figure 10.5). The response function marked S responds most strongly to the shortest wavelengths: blue and violet; the response function marked I responds most strongly to yellow and green, while the response function marked L responds to the longer wavelengths, primarily yellow, orange and red.



These response curves show how the blue and green primary colors were chosen, since they are close to the peaks in the response functions of two of the 3 types of cones. (Also note why some definitions use a primary blue at 425 nm – it is closer to the peak response of one of the types of cones than 460 nm is.) The 3rd primary has a longer wavelength than the peak in the response of the third cone, which is why monochromatic red colors tend to look less bright than the same amount of energy at shorter-wavelength parts of the spectrum. In addition, the response of the “red cone” to red light is only somewhat  greater than the response of the “green cone” to the same wavelength, so that red light often has a hint of yellow in it even when it is really a completely saturated red.

 

The fact that there are three types of cones whose wavelength sensitivities are as shown above explains many of the phenomena that we have described. For example, it is clear why color matching using the three primary colors can work. Since each type of cone reports only its relative excitation level and not the exact wavelength that caused the excitation, it is possible in principle to match the three excitation levels resulting from any incident wavelength using an appropriate combination of the three primaries.

 

It is also clear from the figure why some colors (especially monochromatic blues and greens) cannot be matched using the usual primaries. Any incident wavelength in the range of 460-500 nm will excite the blue cone quite strongly, but will also excite both the green and red cones to some extent. Thus matching colors in this region will need lots of the blue primary. Since the green cone is also excited, some green primary will also be required. Unfortunately, the response of the red cone to this green light is quite significant – much more than the response of the red cone to the incident wavelength. Thus adding green primary to the mix almost always produces too much excitation in the red cone. The only way to match this unintended (but unavoidable) excitation of the red cone is to add some red to the incident wavelength. This conclusion is exactly the same as we came to from considering the color matching horseshoe chart in the previous topics – highly saturated blue/green wavelengths require a “negative” red contribution in order to be matched.

 

Although the response curves of the cones explains color many of the aspects of color matching, it does not explain all of the observed phenomena of color vision. In particular, the fact that the combination of two colors (such as red and green) can produce a third color, whose wavelength is not related in any obvious way to its two parents is not a direct result of the previous discussion. In addition, the interaction between intensity and perceived hue is not explained either. Since the physical response functions of the cones have been verified in many ways, these additional phenomena must be implemented in the processing of the signals from the cones rather than in the hardware of the cones themselves.

 

          While there are only three types of cones, there is some evidence that the processing of these 3 signals can be characterized in terms of differences between the different channels. There are 5 signals in this model: red, green, blue, yellow and intensity. (Based on our previous discussion, the yellow channel is presumably implemented internally as (red + green). The 3 signals that are processed by the brain are the pair-wise differences: red – green, blue – yellow and the overall intensity, which is presumably estimated as the sum of the excitations of the 3 types of cones.  The intensity response function is complicated by other factors. For example, the perceived brightness of an object depends on its surroundings, so that the intensity channel incorporates some kind of spatial differencing.

 

The wavelength values corresponding to these physiological primaries are not quite the same as the primary colors of the previous discussion. The largest discrepancy is the value of the “red” primary, which is closer to what we defined above as magenta – a mixture of red and blue. Therefore, using our previous primary values, the red – green signal is really more like (red + a little blue) – green. Since there is no cone that explicitly responds to yellow, the yellow – blue channel is presumably implemented as (red + green) – blue.

 

          When both difference channels red – green and yellow – blue respond with a 0 value, then the color is some shade of white depending on the value of the intensity channel. The overall response is summarized in the following figure (figure 10.12, page 277).



 

In addition to the differencing in wavelength, there is also some spatial differencing. That is, a color in one area can affect the perception of the color of an adjacent area. That is, a bright yellow area makes an adjacent uncolored area appear blue, etc.

 

Finally, there is an interaction between the perceived color of a scene and its actual average intensity. This tends to work in two ways: (1) since the color processing involves differences between channels, a decrease in the overall illumination does not affect the differences as much, so that we can perceive more or less the same color under different levels of illumination. (2) a change in the intensity of only part of a scene relative to the rest of it does have an effect on the perceived hue – dim orange will look brown, for example.

 

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