Dr. Greenwood received a dual Ph.D. in Integrative Physiology and Neuroscience from the University of Colorado-Boulder in 2005. He then served as a lecturer in the Department of Psychology at the University of Colorado-Denver and Health Scienves Center while completing post-doctoral training in the laboratory of Dr. Monika Fleshner in the Department of Integrative Physiology at the University of Colorado-Boulder. In 2010, he joined the faculty of the Department of Integrative Physiology as an Assistant Research Professor. Dr. Greenwood has been a member of the Society for Neuroscience since 1999.
Dr. Greenwood's research focuses on understanding how physical activity impacts behavioral responses to stress, with particular emphasis on learning and memory, depression, and anxiety behaviors in rodents. Exercise can help prevent many of the deleterious effects of stress on the brain and behavior. Studies in the laboratory implement behavioral neuroscience, molecular, and pharmacological strategies to identify the neurobiological mechanisms underlying exercise-induced stress resistance.
Greenwood BN and Fleshner M. Exercise, stress and central serotonin systems. Exercise and Sports Science Reviews. 2011 Jul:39(3):140-9.
Greenwood BN, Foley TE, Le TV, Strong PV, Loughridge AB, Day HEW, Fleshner M. Long term voluntary exercise is rewarding and produces plastic changes in the mesolimbic reward pathway. Behavioral Brain Research, 2011 Mar 1:217(2):354-62.
Greenwood BN, Strong, PV, Fleshner M. A behavioral analysis of the impact of voluntary physical activity on hippocampal-dependent contextual conditioning. Hippocampus. 2009 Oct: 19(10):988-1001.
Greenwood BN, Strong PV, Brooks L, Fleshner M. Anxiety-like behaviors produced by acute fluoxetine administration in male Fischer 344 rats are prevented by prior exercise. Psychopharmacology, 2008 Aug:199(2):209-22.
Greenwood BN, Foley TE, Day HEW, Burhams D, Brooks L, Campeau S, Fleshner M. Wheel running alters serotonin (5-HT) transporter, 5-HT1A, 50HT1B, and alpha 1b-adrenergic receptor mRNA in the rat raphe nuclei. Biological Psychiatry 2005 Mar 1; 57(5):559-68.
Greenwood BN, Foley TE, Day HE, Campisi J, Hammack SH, Campeau S, Maier SF, Fleshner M. Wheel running prevents learned helplessness/behavioral depression: role of dorsal raphe serotonergic neurons. Journal of Neuroscience 2003 Apr 1; 23(7):2889-98.