Ecology and Evolutionary Biology

The Department of Ecology and Evolutionary Biology (formerly the Department of Environmental, Population, and Organismic Biology) offers two general biology sequences. EBIO 1210 and 1200 are lecture courses intended for EBIO and other science majors. A year of high school chemistry is presumed. Accompanying laboratories (EBIO 1230 and 1240) are also available. EBIO 1030 and 1040 are designed for nonscience majors; an accompanying lab (EBIO 1050) is available.

Students with scores of 4 or 5 on the AP biology test receive 8 hours of credit and are exempt from EBIO 1210–1240. Students who score in the 66th percentile or higher on the CLEP test in biology receive 6 hours of credit and are exempt from EBIO 1210 and 1220. MCDB 1150 and 1151 can substitute for EBIO 1210 and 1230. EBIO majors with transfer credit in biology from other institutions and/or advanced placement credits must consult the EBIO undergraduate advising and resource center for proper placement.

EPOB courses have been renumbered under EBIO and IPHY course numbers. Credit will not be granted for a course taken under its former number and its current number.

EBIO 1030-3. Biology: A Human Approach 1. Lect. Studies the principles of biology and their implications. Central theme is humans and the environment, emphasizing ecology, natural resource conservation, and the interrelatedness of a growing human population. Recommended for nonscience majors. Meets MAPS requirement for natural science: non-lab. Approved for arts and sciences core curriculum: natural science.

EBIO 1040-3. Biology: A Human Approach 2. Lect. Continues EBIO 1030, focusing on the function of the human body, and maintenance of dynamic equilibrium in the internal environment in the face of a continually changing external environment. Discusses factors influencing these homeostatic conditions and how and why they change. Recommended for nonscience majors. Prereq., EBIO 1030. Approved for arts and sciences core curriculum: natural science.

EBIO 1050-1. Biology: A Human Approach Laboratory. One two-hour lab per week. Provides experiments and exercises relating to concepts presented in EBIO 1030 and 1040 Biology: A Human Approach 1 and 2. This course uses animals and/or animal tissues. Recommended for nonscience majors. When taken with EBIO 1030, meets MAPS requirement for natural science: lab. Approved for arts and sciences core curriculum: natural science.

EBIO 1210-3. General Biology 1. Lect. Provides a concentrated introduction to molecular, cellular, genetic, and evolutionary biology. Emphasizes fundamental principles, concepts, facts, and questions. Intended for science majors. Credit not granted for this course and EPOB 2060, 2660 or KAPH 2060. Meets MAPS requirement for natural science. Approved for arts and sciences core curriculum: natural science.

EBIO 1220-3. General Biology 2. Provides a concentrated introduction to organisms, homeostasis, development, behavior, and ecology. Emphasizes fundamental principles, concepts, facts, and questions. Intended for science majors. Prereq., EBIO 1210 or equivalent. Credit not granted for this course and EPOB 2050 or 2650. Approved for arts and sciences core curriculum: natural science.

EBIO 1230-1. General Biology Laboratory 1. One 3-hour lab per week. Consists of experiments and exercises to provide an extension of basic concepts and scientific approaches presented in General Biology 1. Intended for science majors. Prereq. or coreq., EBIO 1210. Credit not granted for this course and EPOB 2060, 2660 or KAPH 2060. Meets MAPS requirement for natural science. Approved for arts and sciences core curriculum: natural science.

EBIO 1240-1. General Biology Laboratory 2. One 3-hour lab per week. Consists of experiments and exercises to provide an extension of basic concepts and scientific approaches presented in General Biology 2. Intended for science majors. Prereq. or coreq., EBIO 1220 or equivalent. Credit not granted for this course and EPOB 2050 or 2650. Approved for arts and sciences core curriculum: natural science.

EBIO 1300 (1-3). Topics in Biological Sciences. Covers special topics in biology for freshmen or nonmajors. Introduces scientific methods and principles in biology, as well as issues of current interest in biology. Does not count toward the major in EBIO.

EBIO 1840 (1-6). Independent Study (Freshman). May be repeated up to 6 total credit hours.

EBIO 2010 (1-3). Environmental Issues and Biology. Lect. Describes how the natural environment is currently stressed by a variety of human actions. Examines the nature of these environmental problems and their impact on living organisms, both human and nonhuman species.

EBIO 2040-4. Principles of Ecology. Lect. and lab. Introduces principles of ecology, emphasizing patterns and processes at various levels of biological organization. Scope global, but examples often from local environment. Laboratory emphasizes techniques of field biology. Uses animals and/or animal tissues. Prereqs., EBIO 1030, 1040, and 1050, or EBIO 1210, 1220, 1230, and 1240. Credit not granted for this course and EPOB 2050 or EPOB 2650 or EPOB 3020 or EBIO 2640.

EBIO 2070-4. Genetics: Molecules to Populations. Lect. and rec. Covers principles of genetics and developmental biology at levels of molecules, cellular organelles, individuals, and populations; asexual and sexual life cycles; and heredity. Recitations allow discussion of genetics problems and implications of genetic principles, and provide demonstrations and simulations of genetic processes. Intended for sophomore majors in EBIO. Prereqs., EBIO 1210, 1220, 1230, and 1240, or EPOB 2050 and 2060. Credit not granted for this course and EBIO 2670.

EBIO 2500-4. Introduction to Horticulture. Lect. and lab. Covers the principles and techniques of plant science applied to cultivated plants. Emphasizes basic plant biology, aspects of the culture environment as variable, and the tools and technology used in culture, regulation, propagation, and protection. Includes a brief survey of the industries related to cultivated plants. Prereqs., EBIO 1210, 1220, 1230, and 1240, or EPOB 2050 and 2060.

EBIO 2590-2. Plants and Society. Acquaints students with the plants that are both essential and desirable to human survival, well-being, and quality of life. Topics include plants and world cultures, food plants, commercial products (beverages, extracts, herbs, and spices, etc.), cosmetics, textiles, wood products, medicinal plants, psychoactive plants, poisonous plants, plant used in horticulture and landscape architecture, wood products, musical instruments, etc.

EBIO 2640-5. Honors Principles of Ecology. Lect., lab, and rec. Introduces principles of ecology, emphasizing patterns and processes at various levels of biological organization. Scope global, but examples often from local environment. Lab emphasizes techniques of field biology. Uses animals and/or animal tissues. Prereqs., EBIO 1210, 1220, 1230, and 1240; or 1030, 1040, and 1050. Credit not granted for this course and EPOB 2050 or 2650 or 3020 or EBIO 2040.

EBIO 2670-5. Honors Genetics: Molecules to Populations. Lect., rec., and co-sem. Covers principles of genetics and developmental biology at levels of molecules, cellular organelles, individuals, and populations; asexual and sexual life cycles; and heredity. Recitations allow discussion of genetics problems and implications of genetic principles, and provide demonstrations and simulations of genetic processes. Honors co-seminar covers specific topics in more depth. Prereqs., EBIO 1210, 120, 1230, and 1240, or EPOB 2050 and 2060. Credit not granted for this course and EBIO 2070.

EBIO 2840 (1-6). Independent Study (Sophomore). May be repeated up to 6 total credit hours.

EBIO 3010 (1-2). Teaching Biology. Provides an opportunity to assist in teaching of specific laboratory section in EBIO under direct faculty supervision. Students must make arrangements with the faculty member responsible for the course in which they plan to assist. May be repeated up to 4 total credit hours.

EBIO 3040-4. Conservation Biology. Applies principles of population ecology, population genetics, biogeography, animal behavior, and paleobiology to the maintenance of biodiversity and natural systems. The resulting theory is then applied to conservation policy and management techniques. Prereq., EBIO 2040 or 2640. Same as ENVS 3040.

EBIO 3080-4. Evolutionary Biology. Lect. and rec. Emphasizes the fundamental evolutionary concepts that provide explanations for the diversification of life on Earth. Specific topics include the evidence for evolution, adaptation by natural selection, speciation, systematics, molecular and genome evolution, and macroevolutionary patterns and processes. Recitations allow students to explore specific topics in more depth and smaller groups. Prereq., EBIO 1210 and 1220 or equivalent. Credit not granted for this course and EBIO 3680 or EPOB 2080 or 2680.

EBIO 3110-3. Population and Community Ecology. Presents principles of ecology that relate to the niche, population growth, metapopulations, population interactions (within and between trophic levels), community structure and development, landscape ecology and species diversity. Prereq., EBIO 2040 or 2640 or equivalent.

EBIO 3170 (3-4). Arctic and Alpine Ecology. Lect. and field trips. Focuses on the biology of arctic and alpine environments, limiting physical factors (such as geomorphology and climatic history), and human interaction with cold-stressed environments, especially the arctic. Prereqs., EBIO 1210 and 1220, or EPOB 2050 and 2060.

EBIO 3175-1. Arctic and Alpine Ecology Lab. Examines alpine ecosystems and treeline relative to global change. Weekend (one-day) field trips into the Rocky Mountains, visits to Denver Museum of Nature and Science, and to CU herbarium/plant lab for experiential learning connected to EBIO 3170. Coreq., EBIO 3170. Pass/fail only.

EBIO 3180-3. Global Ecology. Lect. Involves study of ecological principles and problems at the biosphere level. Presents a worldwide approach to populations, biotic resources, ecologic interactions, land use, deforestation, desertification, species extinctions, pollution, environmental quality, global change, and environmental ethics. Prereqs., EBIO 1210 and 1220 or equivalent. Formerly EPOB 3180. Approved for arts and sciences core curriculum: natural science.

EBIO 3190-3. Tropical Marine Ecology. Lect. Examines the biology and ecology of marine ecosystems, emphasizing those occurring in tropical regions such as coral reefs. Studies how these ecosystems are changing and the future impact of human stress on the marine environment. Prereqs., EBIO 1210, 1220, 1230, and 1240, or EPOB 2050 and 2060. Restricted to sophomores/juniors/seniors. Approved for arts and sciences core curriculum: natural science.

EBIO 3240-4. Animal Behavior. Lect. and lab. Topics include basic concepts and history, methods of study, ethical issues, neurobiology, behavior, the development of behavior, predator-prey relationships, communication, aggression and dominance, mating systems, cognitive ethology, and parental care. When possible, life-history strategies, the evolution of behavior, and behavioral ecology are stressed. Uses animals and animal tissues. Prereqs., EBIO 1210, 1220, 1230, and 1240, or EPOB 2050 and 2060. Restricted to sophomores/ juniors/seniors.

EBIO 3270-3. Ecosystem Ecology. Integrates information from physics (e.g., energetics), chemistry (the behavior of basic elements), and biology (evolutionary traits of species, multiple photosynthetic pathways, etc.) to understand the structure and functioning of ecosystems. Provides the background and necessary information to understand controls on photosynthesis, decomposition, and nutrient cycling across diverse terrestrial and aquatic landscapes. Prereqs., EBIO 1210, 1220, 1230, 1240, and 2040, or EPOB 2050 and 2060, or EBIO 3020.

EBIO 3400-4. Microbiology. Lect. and lab. Surveys distinguishing characteristics of microorganisms based on structural-functional relationships, taxonomy, growth, and physical-chemical agents of control including antibiotics, metabolism, and genetics. Introduces applied microbiology emphasizing infectious diseases, basic concepts of immunology, and microbial ecology. Uses animals and/or animal tissues. Prereqs., EBIO 1210, 1220, 1230, and 1240, or EPOB 2050 and 2060.

EBIO 3470-3. History of Biology. Lect. Surveys major themes in the development of biological theory from ancient times to present, emphasizing complimentary roles of observation, experiment, and technical innovation, and influence of general cultural environment on scientific advance. Prereqs., EBIO 1210 and 1220.

EBIO 3630-4. Parasitology. Lect. and lab. Surveys animal parasites, including life histories; emphasizes parasites of humans. Uses animals and/or animal tissues. Prereqs., EBIO 1210, 1220, 1230, and 1240 or EPOB 2050 and 2060.

EBIO 3680-5. Honors Evolutionary Biology. Lect., rec., and co-sem. Emphasizes the fundamental evolutionary concepts that provide explanation for the diversification of life on Earth. Specific topics include the evidence for evolution, adaptation by natural selection speciation, systematics, molecular and genome evolution, and macroevolutionary patterns and process. Recitations allow students to explore specific topics in more depth and smaller groups. Prereqs., EBIO 1210, 1220, 1230, 1240, 2070 or 2670. Credit not granted for this course and EBIO 3080 or EPOB 2080 or 2680.

EBIO 3770-4. Animal Diversity: Vertebrates. Lect. and lab. Provides a broad overview of the biology of vertebrates: evolution and systematics, morphology, physiology, behavior, ecology, and biogeography. Laboratories focus on the diversity of vertebrates, including adaptations and diagnostic features of major groups. Uses animals and/or animal tissues. Prereqs., EBIO 1210, 1220, 1230, and 1240, or equivalent.

EBIO 3840 (1-6). Independent Study (Junior). May be repeated up to 6 total credit hours.

EBIO 3850-4. Animal Diversity: Invertebrates. Lect. and lab. Offers a broad study of the biology of the most diverse group of organisms on Earth. Areas include ecology, physiology, evolution and morphology of aquatic and terrestrial forms. This course uses animals and/or animal tissues. Prereqs., EBIO 1210, 1220, 1230, and 1240, or equivalent, or EPOB 2050 and 2060.

EBIO 3870 (1-6). Independent Research (Junior). May be repeated up to 6 total credit hours.

EBIO 3930 (1-6). Internship. Provides an academically supervised opportunity for upper-division students to work in public or private organizations. Projects are usually related to students’ career goals. Each project has both academic and work components. May be repeated up to 6 total credit hours. Pass/fail only.

EBIO 3940-3. Argument in Scientific Writing. Emphasizes argumentative strategies used in scientific writing. Reviews essential writing skills to prepare students for academic and professional communication. Restricted to juniors and seniors. No biology credit for majors in EBIO. Approved for arts and sciences core curriculum: written communication.

EBIO 3980-1. Seminar: Introduction to EBIO Honors. Presents an introduction to the departmental Honors program. Consists of a lecture component on Honors research, thesis, and defense, as well as a seminar component where students present the findings of their library research, conducted under guidance of a faculty mentor, and hear presentations by graduating Honors candidates on their thesis research. Prereq., 3.2 or better GPA, declared EBIO major, and approval by departmental Honors program.

EBIO 3990 (1-3). Introduction to EBIO Honors. Provides an introduction to the departmental Honors program. Consists of individual library research on a potential Honors thesis topic under the guidance of a faculty mentor. Prereqs., 3.2 or better GPA, declared EBIO major, and approval by departmental Honors committee.

EBIO 4020-3. Stream Biology. Offers a geological, physical, chemical, and biological study of flowing water with special reference to streams and rivers as ecosystems. A laboratory course is offered (see EBIO 4150). Prereqs., EBIO 1210, 1220, 1230, 1240, and 2040, or EPOB 2050 and 2060. Same as EBIO 5020.

EBIO 4030-3. Limnology. Examines the ecology of inland waters, including a detailed consideration of physical, chemical, and biological properties of freshwater ecosystems: origins and major characteristics of lakes and streams, survey of chemical and nutrient cycles in freshwater habitats, and survey of biotic composition of freshwater environments. Important themes in modern freshwater ecology are considered, including energy flow, trophic structure, eutrophication, and management of freshwater ecosystems. Prereqs., EBIO 1210 and 1220. Same as EBIO 5030.

EBIO 4060-3. Landscape Ecology. Studies distributional patterns of communities and ecosystems, ecological processes that affect those patterns, and changes in pattern and process over time. Consideration of spatial and temporal scales in ecological analyses is required to understand and predict response to broad-scale environmental change. Prereqs., EBIO 1210, 1220, 1230, 1240, and EBIO 2040 or 2640, or EPOB 2050 and 2060. Same as EBIO 5070.

EBIO 4090-2. Coral Reef Ecology. Includes one week of lectures in Boulder and one week of field studies on one of the most complex and beautiful ecosystems in the world, the Caribbean reefs at Cozumel, Mexico. Two week, fall-semester course beginning after Christmas. Prereqs., EBIO 2040 and SCUBA certification. Restricted to sophomores/juniors/ seniors.

EBIO 4100-3. Advanced Ecology. Emphasizes specific aspects of ecology based on specialties of faculty. One or more courses are offered most semesters. Topics have included dynamics of mountain ecosystems, tundra ecology, ethnoecology, population dynamics, tropical and insular biology, ecology of fishes, quantitative plant ecology, and arctic and alpine environments. May use animals and/or animal tissues. May be repeated twice provided the topics vary. Prereqs., EBIO 1210, 1220, 1230, 1240, and EBIO 2040 or 2640, or EPOB 2050 and 2060. Same as EBIO 5100.

EBIO 4110-3. Advanced Ecology. See EBIO 4100 for description. May use animals and/or animal tissues. May be repeated up to 9 total credit hours. Prereqs., EBIO 1210, 1220, 1230, 1240, and EBIO 2040 or 2640, or EPOB 2050 and 2060. Same as EBIO 5110.

EBIO 4120 (2-4). Advanced Ecology. See EBIO 4100 for description. May use animals and/or animal tissues. Prereq., EBIO 1210, 1220, 1230, 1240, and EBIO 2040 or 2640, or EPOB 2050 and 2060. Same as EBIO 5120.

EBIO 4140-3. Plant Ecology. Examines the relationships between plants and their physical and biological environments, encompassing physiology, competition, plant-soil and plant-animal interactions, population dynamics, diversity, and influence on ecosystem function. Prereqs., EBIO 1210, 1220, 1230, and 1240.

EBIO 4150 (1-2). Techniques in Ecology. Emphasizes application of modern ecological techniques, such as stream biology, aquatic biology, environmental measurement and control, and techniques in geoecology. May be repeated up to 7 total credit hours. Prereqs., EBIO 1210, 1220, 1230, 1240, and EBIO 2040 or 2640, or EPOB 2050 and 2060. Same as EBIO 5150.

EBIO 4160-3. Introduction to Biogeochemistry. Covers fundamentals of biogeochemical cycling, emphasizing water, carbon, and nutrient dynamics in terrestrial ecosystems; chemical interactions of atmosphere, biosphere, lithosphere, and hydrosphere; and natural and human-managed environments. Prereqs., GEOL 3320 or EBIO 3270, and CHEM 1011 or higher. Same as ENVS 4160 and GEOL 4160.

EBIO 4175-3. The Scientific Basis for Ecosystem Management of Public Lands. An advanced field ecology course emphasizing measurements, statistical procedures, and biotic data information management relevant to land management issues. Develops concepts of adaptive ecosystem management using ongoing field studies on public land in the Colorado Front Range. Prereqs., EBIO 3270 and 3500.

EBIO 4180-3. Ecological Perspectives on Global Change. Discusses evolutionary and recent geological history of modern environmental problems, using natural changes in climate, biotic diversity, drought, desertification, flood, forest destruction, etc., to show the range and frequency of such events as a perspective on modern reports. Prereq., minimum 14 hours of EBIO course work, including EBIO 2040, EPOB 2050 or 3020.

EBIO 4290-3. Molecular Systematics and Evolution. Reviews principles and theoretical foundations of molecular evolution and molecular systematics. Emphasizes hypothesis testing and using various methods of analysis for interpreting molecular variation in nature. Prereq., EBIO 3080 or instructor consent. Same as EBIO 5290.

EBIO 4350 (1-4). Biological Field Studies. Stresses broad areas of biology and employs field approaches. Uses animals and/or animal tissues. May be repeated up to 4 total credit hours. Prereqs., EBIO 1210, 1220, 1230, and 1240, or EPOB 2050 and 2060. Same as EBIO 5350.

EBIO 4410-4. Biometry. Lect. and lab. Offers a demanding, problems-oriented methods course in statistical inference procedures, assumptions, limitations, and applications emphasizing techniques appropriate to realistic biological problems. Includes data file management using interactive computing techniques. Prereqs., EBIO 1210, 1220, 1230, and 1240, or EPOB 2050 and 2060. Restricted to juniors/seniors. Same as EBIO 5410.

EBIO 4460 (1-5). Special Topics. Familiarizes students with specialized areas of biology. May be repeated up to 9 total credit hours. Prereqs., EBIO 1210, 1220, 1230, and 1240, or equivalent. Same as EBIO 5460.

EBIO 4500-4. Plant Biodiversity and Evolution. Lect. and lab. Surveys plant types emphasizing diagnostic features of plants in general and major taxa in particular. Focuses on identity, morphology, anatomy, reproduction, ecology, geography, evolution, fossil record, and economic use of taxa. Prereqs., EBIO 1210, 1220, 1230, and 1240, or EPOB 2050 and 2060. Formerly EBIO 3500.

EBIO 4510-4. Plant Anatomy and Development. Lect. and lab. Introduces structures of seed plants, especially angiosperms, and developmental history of these structures. Studies cell types, and their location and function in plant tissues and organs. The laboratory provides an opportunity to examine plant tissues and to prepare tissues for examination by the light microscope. Stresses role of plant structures in the living plant. Prereqs., EBIO 1210, 1220, 1230, and 1240, or EPOB 2050 and 2060. Formerly EBIO 3510.

EBIO 4520-3. Plant Systematics. Lect. and lab. Studies the principles and techniques of modern systematics of organisms, illustrated with examples from the plant kingdom, usually the angiosperms. Framework of course is evolutionary and ecological, as well as taxonomic. Prereq., EBIO 2070. Restricted to sophomores/juniors/seniors. Formerly EBIO 3520.

EBIO 4530-4. Functional Plant Biology. Lect. and lab. Explores mechanisms of plant functioning and how such functioning relates to the performance of the plant under different environmental conditions. Phenomena include water relations, growth and development, and metabolic processes including photosynthesis, respiration, and responses to stress. Prereqs., EBIO 1210, 1220, 1230, and 1240, or equivalent. Restricted to sophomores/juniors/seniors. Formerly EBIO 3530.

EBIO 4550-4. Plant Eco-Evo-Devo. Explores the fundamental principles of plant form from the perspectives of ecological function, evolutionary origin, and developmental dynamics. Students are presented with conceptual and analytical tools to interpret the vast diversity of growth form-function relationships that exist among plants. Laboratory sessions apply concepts presented in the lecture and students will engage in original research using light and scanning electron microscopy. Prereqs., EBIO 1210, 1220, 1230, or 1240.

EBIO 4570-3. Advanced Plant Physiology. Evaluates critically various concepts underlying the functioning of plants, including current controversial topics. Emphasizes the responses of plants to various environmental factors. Prereq., EBIO 3530. Restricted to sophomores/juniors/seniors. Same as EBIO 5570.

EBIO 4630 (2-6). Field Techniques in Environmental Science. Applies field and laboratory methods for assessing the abiotic and biotic environment. Emphasizes field techniques in climatology, surveying soils, hydrology, geomorphology, plant and animal ecology, and environmental law. Evaluation by written module reports and maps. This course may use animals and/or animal tissues. Prereqs., EBIO 2050 and instructor consent. Same as ENVS 4630.

EBIO 4640 (2-4). Plant Field Studies. Includes field-oriented courses offered at irregular intervals during the academic year or during summer sessions. May be repeated up to 7 total credit hours. Prereq., EBIO 2040 and 2640 or equivalent.

EBIO 4660-4. Insect Biology. Lect. and lab. Introduction to evolution, ecology, physiology, and behavior of insects. Emphasizes how insects have solved problems, such as maintaining water balance or finding food, that are shared by all animals but for which there may be unique solutions among the insects. Agricultural and human health problems relative to entomology are discussed. Uses animals and/or animal tissues. Prereqs., EBIO 1210, 1220, 1230, and 1240, or EPOB 2050 and 2060. Same as EBIO 5660.

EBIO 4740-3. Biology of Amphibians and Reptiles. Comparative morphology, taxonomy, ecology, behavior, and geographic distribution of amphibians and reptiles. This course uses animals and animal tissue. Prereqs., EBIO 1210, 1220, 1230, and 1240, or EPOB 2050 and 2060. Same as EBIO 5740 and PSYC 4740.

EBIO 4750-4. Ornithology. Lect., lab, and field trips. Presents origin, evolution, ecology, physical and behavioral characteristics, and taxonomy of orders and families of birds of North America; field work with local species emphasizing avian ecology. Uses animals and/or animal tissues. Prereq., EBIO 1210, 1220, 1230, and 1240, or EPOB 2050 and 2060. Same as EBIO 5750.

EBIO 4760-4. Mammalogy. Lect., lab, and field studies. Discusses origin, evolution and adaptation, geographic distribution, ecology, and taxonomy of mammals; field and laboratory study of Coloradan species. Uses animals and/or animal tissues. Prereq., EBIO 1210, 1220 and 2040 or 2640. Same as EBIO 5760.

EBIO 4795-3. Museum Field Methods/Zoology and Botany. Class covers research and field methods for biological disciplines associated with natural history museums: vertebrates, invertebrates, and plants. Emphasis is on field research techniques: observations, sampling, collection and preservation methods, and comparisons among elevation zones. Includes 5 field labs, 2 weekend trips, 5 lab practica, experience with several taxonomic experts, and individual research projects. Same as EBIO 5795, MUSM 4795 and ENVS 4795.

EBIO 4800-3. Critical Thinking in Biology. Lect. and discussion. Explores controversial issues, historical themes, or emerging developments in biology. Consult the EBIO Undergraduate Advising Center for current listings. Different course sections on different topics may be repeated up to 12 total credit hours. Prereq., minimum of 14 hours of EBIO course work. Restricted to sophomores/juniors/ seniors. Same as EBIO 5800.

EBIO 4820-3. Scientific Honors Thesis Writing Seminar. Provides advanced instruction in writing the scientific honors theses, emphasizing interpreting, analyzing, and arguing scientific data. Focuses on essential communication skills necessary for effective presentation of the thesis to a wide variety of audiences.

EBIO 4840 (1-6). Independent Study (Senior). May be repeated up to 6 total credit hours.

EBIO 4860 (1-2). Critical Thinking in Biology: Lab.

EBIO 4870 (1-6). Independent Research (Senior). May be repeated up to 6 total credit hours.

EBIO 4980-1. Seminar: EBIO Honors Thesis. To be taken during the final academic year prior to graduation. Consists of a lecture component on honors thesis writing and defense, as well as a seminar component where honors candidates present their thesis research in a practice defense talk. Prereq., 3.30 or better GPA, declared EBIO major, and approval by departmental honors committee.

EBIO 4990 (1-3). EBIO Honors Thesis. To be taken during the final academic year prior to graduation. Consists of the final phase of honors research and thesis preparation under the guidance of a faculty mentor. Prereq., 3.30 or better GPA, declared EBIO major, and approval by departmental honors committee.

EBIO 5000-1. EBIO Colloquia. All first year EBIO graduate students are required to attend the EBIO Colloquia Series. Speakers from around the world and within the department cover topics in all areas of biology. May be repeated up to 2 total credit hours.

EBIO 5020-3. Stream Biology. Same as EBIO 4020.

EBIO 5030-3. Limnology. Same as EBIO 4030.

EBIO 5060-3. Landscape Ecology. Same as EBIO 4060.

EBIO 5100-3. Advanced Ecology. Same as EBIO 4100.

EBIO 5110-3. Advanced Ecology. Same as EBIO 4110.

EBIO 5120 (2-4). Advanced Ecology. Same as EBIO 4120.

EBIO 5150 (1-2). Techniques in Ecology. Same as EBIO 4150.

EBIO 5240-3. Advanced Topics in Animal Behavior. Covers special areas of ethology such as sociobiology, animal communication, cognitive ethology, human ethology, moral and ethical issues. Prereq., EBIO 3240.

EBIO 5290-3. Molecular Systematics and Evolution. Same as EBIO 4290.

EBIO 5310-3. Ecology and Conservation. Reviews current research activities in the various fields of environmental biology, including population, community, behavioral, ecosystem, and landscape ecology, plus conservation biology and global change. Prereq., graduate standing.

EBIO 5320-3. Current Topics in Evolutionary Biology. Examines six major themes on contemporary evolutionary research: population genetics, natural selection and adaptation, molecular evolution, evolution and development, phylogenetic systematics, and macroevolution. Emphasizes recent primary literature and sophisticated mastery. Prereq., graduate standing in EBIO. Formerly EPOB 5320.

EBIO 5350 (1-4). Biological Field Studies. Same as EBIO 4350.

EBIO 5410-4. Biometry. Same as EBIO 4410.

EBIO 5460 (1-5). Special Topics. Same as EBIO 4460.

EBIO 5570-3. Advanced Plant Physiology. Same as EBIO 4570.

EBIO 5660-4. Insect Biology. Same as EBIO 4660.

EBIO 5740-3. Biology of Amphibians and Reptiles. Same as EBIO 4740 and PSYC 5740.

EBIO 5750-4. Ornithology. Same as EBIO 4750.

EBIO 5760-4. Mammalogy. Same as EBIO 4760.

EBIO 5795-3. Museum Field Methods/Zoology and Botany. Same as EBIO 4795 and MUSM 5795.

EBIO 5800-3. Critical Thinking in Biology. Same as EBIO 4800.

EBIO 5820 (2-3). Graduate Writing Seminar. Enhances writing proficiency, using graduate writing projects to implement the course concepts. Offers understanding of conventions and strategies used in scientific writing to prepare students for academic and professional communication. Prereqs., graduate standing and basic proficiency in English as a written language.

EBIO 5840 (1-6). Independent Study (Master’s Level). Instructor consent required. May be repeated up to 6 total credit hours.

EBIO 6000-1. Seminar: Introduction to Biological Research. Discusses areas of biological research represented in EBIO. Required of all first-year graduate students in EBIO.

EBIO 6100 (1-3). Seminar in Environmental Biology. Instructor consent required. Restricted to graduate students. May be repeated up to 7 total credit hours. Same as EBIO 6120.

EBIO 6120 (1-3). Seminar in Environmental Biology. Same as EBIO 6100.

EBIO 6200 (1-3). Seminar in Population Biology. May be repeated up to 7 total credit hours. Same as EBIO 6210.

EBIO 6210 (1-3). Seminar in Population Biology. May be repeated up to 7 total credit hours. Same as EBIO 6200.

EBIO 6300 (1-3). Seminar in Organismic Biology. May be repeated up to 7 total credit hours.

EBIO 6330-2. Seminar in Organismic Biology. May be repeated up to 7 total credit hours.

EBIO 6440-2. Remote Sensing Field Methods. Explores theory and practical field measurements for validation of airborne and spaceborne spectral image acquisition. Emphasizes radiative scattering properties of soil, vegetation, cryosphere, and atmosphere. Characterization and calibration of instrumentation used to measure these properties. Prereq., GEOL/GEOG 4093/5093. Recommended prereq., GEOL 5240. Same as GEOG 6443.

EBIO 6840 (1-7). Independent Research (Master’s Level). Instructor consent required. May be repeated up to 7 total credit hours.

EBIO 6940 (1-6). Master’s Degree Candidate: Plan II. Prereq., instructor consent.

EBIO 6950 (1-6). Master’s Thesis. May be repeated up to 7 total credit hours. Prereq., instructor consent.

EBIO 7840 (1-6). Independent Study (Doctoral Level). May be repeated up to 6 total credit hours.

EBIO 8840 (1-6). Independent Research (Doctoral Level). May be repeated up to 6 total credit hours.

EBIO 8990 (1-10). Doctoral Dissertation. Prereq., instructor consent.

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